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APS 323 'Social Insects' course   

Co-ordinator: Professor F. Ratnieks

The estimated 20,000 species of eusocial insects (termites, ants and some bees and wasps) are of great economic and ecological significance. They are also model organisms for studying many fundamental questions in biology. The course is divided into three sections. The first section provides background information on social insect biology. The other two sections focus on two important general questions in biology for which insect societies are excellent model systems, and which are studied in Prof. Ratnieks’ laboratory.

1) Background to social insects. What is eusociality and why study social insects; the behavioural and morphological specializations of individuals (queens, workers, males) to eusocial life; division of labour; colony life cycles; taxonomic diversity of social insects etc.

2) Reproductive conflicts and conflict resolution within insect societies. That is, how the non-clonal kin structure of insect societies leads to conflicts among individuals over reproduction and how these conflicts can be resolved. This section of the course will also introduce students to inclusive fitness theory.

3) How insect societies organize themselves. That is, how adaptive colony-level organization, such as effective foraging, arises from the actions of individual workers.

Lectures will be by Prof. Ratnieks, with one or two guest lectures and a video.

Aims and Objectives

Lecture Content (download the 2007 Lectures and Handouts, compressed file .rar)

1. Introduction to social insects
Lectures 1 & 2 Why study social insects?
Handout
Slideshow
Are You Being Served?
Beekeeping
Big Businesses
Conflict in the Bee Hive
Eusociality Primer
Garbage Disposal in Leafcutter Ants
How Bees Make Honey
Outsmarted by Ants
Policing Insect Societies
Lecture 3 The individual
Handout
Slideshow
Lecture 4 The society
Handout
Slideshow
   
2. Reproductive conflicts and conflict resolution
Lecture 5 Biodiversity and Systematics
Handout
Slideshow
Lecture 6 Division of Labour
Handout
Ant Allometry
Slideshow
Lecture 7 Kin Value
Handout (additional material for those interested)
Slideshow
Lecture 8 Kin Structure and Relatedness
Handout
Slideshow
Lecture 9 Sex Allocation
Handout
Slideshow
Lecture 10 Male Production
Handout
Slideshow
Lecture 11 Caste Fate and Synthesis
Handout
Slideshow
Policing Insect Societies
Conflict Resolution in Insect Societies
   
Section 3. How insect societies organize themselves
Lecture 12 Principles of Self-organisation
Handout
Slideshow
Lecture 13 Honey Bee Foraging: How Colonies Track Food
Handout
Slideshow
Lecture 14 Pharaoh's Ant Foraging: Sophisticated Networks
Handout
Slideshow
Trail Geometry
Outsmarted by ants
Negative Pheromone
Lecture 15 Task Partitioning in Insect Societies
Handout
Slideshow
Guest Lecture
Dr. William Hughes
Defending the Fortress: Host-Parasite Interactions
Handout
Slideshow
   
Additional Materials

Coursework question 1
Coursework template, Pamphlet examples (compressed file .rar)
Notes on video "For Queen and Colony"

PAST EXAMS

Final exam 2004

Final exam 2003

Interesting Articles (compressed file .rar)

 

Update: 22 Feb. 2007

Prof. Francis Ratnieks
Department of Animal and Plant Sciences,
Sheffield University,Western Bank, Sheffield S10 2TN, UK
tel: 0114 2220070 (FLWR office); 01142 2220149 (lab); 01142 220147 (FLWR lab office);
fax: 0114 2220002
e-mail: F.Ratnieks@Sheffield.ac.uk
(when dialing from ouside UK the country code is 44; also delete the 0 in the 0114 city code)

 

Where do wasps go in winter?